Kirke considers herself first and foremost an artist. She began acting through roles she accepted for a number of her friends. Dunham asked Kirke to take on a supporting role in her debut film Tiny Furniture. Kirke and a number of other friends were called upon as a favour to Dunham since there was not enough money to pay professional actors. Although the film turned out to be profitable, Kirke received no payment. Kirke reunited with Dunham in the HBO series Girls, appearing in all six seasons from 2011 to 2017 as the character Jessa Johansson. Kirke next had supporting roles in the dark comedy films Ava's Posessions and The Little Hours. In 2018, Kirke starred alongside her real-life sister Lola Kirke in Emma Forrest's film Untogether. The film premiered at the 2018 Tribeca Film Festival.
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